Reference:

Ma G-H, Bunn E (2007) Embryology and pollination trials support dichogamy in Santalum album L. Sandalwood Research Newsletter 22, 1-4.

Abstract:

Embryo development and pollination trials were studied in Santalum album L. The formation of the male (microspore) and female (megaspore) tissues in the same flower were synchronized during the early stages of flower-bud development. However, at anthesis when pollen was mature, the megaspore had developed only to the stage of a 1-2 nucleate embryo sac. The development from 2-nucleate embryo sac to matured 8-nucleate embryo sac lasted up to 10 days. These results indicate that the flower of S. album is dichogamous where the pollen matures before the embryo sac. Following fertilisation of the ovule the endosperm developed prior to division of the zygotic embryo, and 1-3 embryos and endosperms were formed in the same fruit. Seed-set resulting from open pollination was less than 3%. No seed set was observed when inflorescences were covered with a bag; however artificial cross pollination increased fruit set to 14%. Mature seed usually germinated to produce one seedling, but two- and three-seedlings from one seed were also observed at low frequency.

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